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class=\"wp-block-paragraph\">\u8840\u6d41\u306e\u4f4e\u4e0b\u306f\u6bdb\u6839\u3078\u306e\u9178\u7d20\u3084\u6804\u990a\u306e\u4f9b\u7d66\u3092\u59a8\u3052\u3001\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u306e\u6a5f\u80fd\u3092\u4f4e\u4e0b\u3055\u305b\u308b\u4e00\u56e0\u3068\u306a\u308a\u307e\u3059\u3002\u66f4\u5e74\u671f\u3092\u8fce\u3048\u305f\u5973\u6027\u306b\u300c\u6025\u306b\u767d\u9aea\u304c\u5897\u3048\u305f\u300d\u3068\u611f\u3058\u308b\u65b9\u304c\u591a\u3044\u306e\u306f\u3001\u3053\u3046\u3057\u305f\u8907\u5408\u7684\u306a\u30db\u30eb\u30e2\u30f3\u5909\u52d5\u304c\u80cc\u666f\u306b\u3042\u308b\u305f\u3081\u3067\u3059\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u7532\u72b6\u817a\u6a5f\u80fd\u306e\u4f4e\u4e0b\u3082\u767d\u9aea\u306b\u76f4\u7d50\u3059\u308b<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\u7532\u72b6\u817a\u30db\u30eb\u30e2\u30f3\u306f\u4ee3\u8b1d\u5168\u822c\u3092\u30b3\u30f3\u30c8\u30ed\u30fc\u30eb\u3057\u3001\u6bdb\u5305\u5185\u3067\u306e\u30e1\u30e9\u30cb\u30f3\u5408\u6210\u306b\u3082\u95a2\u4e0e\u3057\u3066\u3044\u307e\u3059\u3002\u7532\u72b6\u817a\u6a5f\u80fd\u4f4e\u4e0b\u75c7\u3067\u306f\u8272\u7d20\u306e\u751f\u6210\u304c\u6ede\u308a\u3001\u767d\u9aea\u304c\u9032\u884c\u3059\u308b\u5834\u5408\u304c\u3042\u308a\u307e\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">40\u4ee3\u4ee5\u964d\u306e\u5973\u6027\u306f\u7532\u72b6\u817a\u306e\u6a5f\u80fd\u7570\u5e38\u3092\u8d77\u3053\u3057\u3084\u3059\u3044\u305f\u3081\u3001\u767d\u9aea\u304c\u6025\u306b\u5897\u3048\u305f\u3068\u304d\u306f\u7532\u72b6\u817a\u306e\u691c\u67fb\u3092\u53d7\u3051\u308b\u306e\u3082\u9078\u629e\u80a2\u306e\u4e00\u3064\u3067\u3059\u3002<\/p>\n\n\n\n<figure class=\"wp-block-table\"><table class=\"has-fixed-layout\"><thead><tr><th>\u30db\u30eb\u30e2\u30f3\u306e\u7a2e\u985e<\/th><th>\u767d\u9aea\u3078\u306e\u5f71\u97ff<\/th><th>\u5909\u52d5\u304c\u8d77\u304d\u3084\u3059\u3044\u6642\u671f<\/th><\/tr><\/thead><tbody><tr><td>\u30a8\u30b9\u30c8\u30ed\u30b2\u30f3<\/td><td>\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u4fdd\u8b77\u529b\u304c\u4f4e\u4e0b<\/td><td>40\u4ee3\u5f8c\u534a\u301c50\u4ee3<\/td><\/tr><tr><td>\u7532\u72b6\u817a\u30db\u30eb\u30e2\u30f3<\/td><td>\u30e1\u30e9\u30cb\u30f3\u5408\u6210\u304c\u6ede\u308b<\/td><td>40\u4ee3\u301c<\/td><\/tr><tr><td>\u30b3\u30eb\u30c1\u30be\u30fc\u30eb<\/td><td>\u30b9\u30c8\u30ec\u30b9\u53cd\u5fdc\u3067\u5e79\u7d30\u80de\u3092\u6d88\u8017<\/td><td>\u5168\u5e74\u4ee3<\/td><\/tr><\/tbody><\/table><\/figure>\n\n\n\n<h2 class=\"wp-block-heading\">\u9178\u5316\u30b9\u30c8\u30ec\u30b9\u304c\u6bdb\u6839\u306e\u8272\u7d20\u7d30\u80de\u3092\u8ffd\u3044\u8a70\u3081\u308b<\/h2>\n\n\n\n<p class=\"wp-block-paragraph\">\u767d\u9aea\u306e\u9032\u884c\u306b\u306f\u6d3b\u6027\u9178\u7d20\u306b\u3088\u308b\u9178\u5316\u30b9\u30c8\u30ec\u30b9\u304c\u6df1\u304f\u95a2\u4e0e\u3057\u3066\u3044\u307e\u3059\u3002\u6bdb\u5305\u5185\u306b\u904e\u9178\u5316\u6c34\u7d20\u304c\u84c4\u7a4d\u3059\u308b\u3068\u3001\u30e1\u30e9\u30cb\u30f3\u3092\u5408\u6210\u3059\u308b\u9175\u7d20\u30c1\u30ed\u30b7\u30ca\u30fc\u30bc\u304c\u6a5f\u80fd\u4e0d\u5168\u3092\u8d77\u3053\u3057\u3001\u9aea\u306b\u8272\u3092\u3064\u3051\u3089\u308c\u306a\u304f\u306a\u308a\u307e\u3059\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u904e\u9178\u5316\u6c34\u7d20\u304c\u6bdb\u5305\u5185\u306b\u84c4\u7a4d\u3059\u308b\u3068\u3069\u3046\u306a\u308b\u304b<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">2009\u5e74\u306bFASEB Journal\u8a8c\u306b\u767a\u8868\u3055\u308c\u305f\u7814\u7a76\u3067\u306f\u3001\u767d\u9aea\u306e\u6bdb\u5e79\u5185\u90e8\u306b\u30df\u30ea\u30e2\u30eb\u5358\u4f4d\u306e\u904e\u9178\u5316\u6c34\u7d20\u304c\u84c4\u7a4d\u3057\u3066\u3044\u308b\u3053\u3068\u304c\u521d\u3081\u3066\u78ba\u8a8d\u3055\u308c\u307e\u3057\u305f\u3002\u904e\u9178\u5316\u6c34\u7d20\u306f\u30e1\u30e9\u30cb\u30f3\u5408\u6210\u306e\u9375\u3092\u63e1\u308b\u30c1\u30ed\u30b7\u30ca\u30fc\u30bc\u306e\u6d3b\u6027\u90e8\u4f4d\u3092\u9178\u5316\u3057\u3001\u9175\u7d20\u306e\u50cd\u304d\u3092\u6b62\u3081\u3066\u3057\u307e\u3044\u307e\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u3044\u308f\u3070\u9aea\u304c\u5185\u5074\u304b\u3089\u300c\u8131\u8272\u300d\u3055\u308c\u3066\u3044\u308b\u72b6\u614b\u3067\u3042\u308a\u3001\u3053\u308c\u306f\u52a0\u9f62\u306b\u4f34\u3046\u6297\u9178\u5316\u9632\u5fa1\u80fd\u529b\u306e\u4f4e\u4e0b\u3068\u76f4\u7d50\u3057\u3066\u3044\u307e\u3059\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u6297\u9178\u5316\u9175\u7d20\u30ab\u30bf\u30e9\u30fc\u30bc\u306e\u6e1b\u5c11\u304c\u767d\u9aea\u3092\u62db\u304f<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\u30ab\u30bf\u30e9\u30fc\u30bc\u306f\u904e\u9178\u5316\u6c34\u7d20\u3092\u6c34\u3068\u9178\u7d20\u306b\u5206\u89e3\u3059\u308b\u6297\u9178\u5316\u9175\u7d20\u3067\u3059\u3002\u82e5\u3044\u9803\u306f\u3053\u306e\u9175\u7d20\u304c\u6bdb\u5305\u3092\u9178\u5316\u30c0\u30e1\u30fc\u30b8\u304b\u3089\u5b88\u3063\u3066\u3044\u307e\u3059\u304c\u3001\u52a0\u9f62\u3068\u3068\u3082\u306b\u30ab\u30bf\u30e9\u30fc\u30bc\u306e\u767a\u73fe\u91cf\u306f\u8457\u3057\u304f\u4f4e\u4e0b\u3057\u307e\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u30ab\u30bf\u30e9\u30fc\u30bc\u304c\u6e1b\u308b\u3068\u904e\u9178\u5316\u6c34\u7d20\u306e\u51e6\u7406\u304c\u8ffd\u3044\u3064\u304b\u306a\u304f\u306a\u308a\u3001\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u304c\u640d\u50b7\u3092\u53d7\u3051\u3066\u8272\u7d20\u3092\u4f5c\u308c\u306a\u304f\u306a\u308a\u307e\u3059\u3002\u30e1\u30c1\u30aa\u30cb\u30f3\u30b9\u30eb\u30db\u30ad\u30b7\u30c9\u9084\u5143\u9175\u7d20\u306e\u6e1b\u5c11\u3082\u52a0\u308f\u3063\u3066\u3001\u9175\u7d20\u306e\u4fee\u5fa9\u80fd\u529b\u81ea\u4f53\u304c\u5931\u308f\u308c\u3066\u3044\u304f\u306e\u3067\u3059\u3002<\/p>\n\n\n\n<h4 class=\"wp-block-heading\">\u9178\u5316\u30b9\u30c8\u30ec\u30b9\u306b\u95a2\u308f\u308b\u4e3b\u306a\u7269\u8cea\u3068\u50cd\u304d<\/h4>\n\n\n\n<figure class=\"wp-block-table\"><table class=\"has-fixed-layout\"><thead><tr><th>\u7269\u8cea\u30fb\u9175\u7d20<\/th><th>\u6b63\u5e38\u6642\u306e\u50cd\u304d<\/th><th>\u52a0\u9f62\u306b\u3088\u308b\u5909\u5316<\/th><\/tr><\/thead><tbody><tr><td>\u30ab\u30bf\u30e9\u30fc\u30bc<\/td><td>\u904e\u9178\u5316\u6c34\u7d20\u3092\u5206\u89e3<\/td><td>\u767a\u73fe\u91cf\u304c\u4f4e\u4e0b<\/td><\/tr><tr><td>\u904e\u9178\u5316\u6c34\u7d20<\/td><td>\u5c11\u91cf\u306a\u3089\u751f\u4f53\u5185\u3067\u81ea\u7136\u767a\u751f<\/td><td>\u84c4\u7a4d\u3057\u3066\u6bdb\u5305\u3092\u9178\u5316<\/td><\/tr><tr><td>\u30c1\u30ed\u30b7\u30ca\u30fc\u30bc<\/td><td>\u30e1\u30e9\u30cb\u30f3\u5408\u6210\u3092\u89e6\u5a92<\/td><td>\u9178\u5316\u3067\u6a5f\u80fd\u304c\u505c\u6b62<\/td><\/tr><\/tbody><\/table><\/figure>\n\n\n\n<h3 class=\"wp-block-heading\">\u7d2b\u5916\u7dda\u3068\u55ab\u7159\u304c\u9178\u5316\u30c0\u30e1\u30fc\u30b8\u3092\u5897\u5e45\u3059\u308b<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\u9178\u5316\u30b9\u30c8\u30ec\u30b9\u306f\u52a0\u9f62\u3060\u3051\u3067\u751f\u3058\u308b\u3082\u306e\u3067\u306f\u3042\u308a\u307e\u305b\u3093\u3002\u7d2b\u5916\u7dda\u3092\u9577\u6642\u9593\u6d74\u3073\u308b\u3068\u6bdb\u5305\u5185\u306e\u6d3b\u6027\u9178\u7d20\u304c\u5897\u52a0\u3057\u3001\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u3078\u306e\u8ca0\u62c5\u304c\u9ad8\u307e\u308a\u307e\u3059\u3002\u5e3d\u5b50\u3084\u65e5\u5098\u3067\u982d\u76ae\u3092\u5b88\u308b\u3053\u3068\u304c\u767d\u9aea\u5bfe\u7b56\u306b\u3082\u3064\u306a\u304c\u308b\u306e\u3067\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u55ab\u7159\u3082\u6d3b\u6027\u9178\u7d20\u3092\u5927\u91cf\u306b\u767a\u751f\u3055\u305b\u308b\u539f\u56e0\u306e\u4e00\u3064\u3067\u3059\u3002\u8907\u6570\u306e\u7814\u7a76\u3067\u55ab\u7159\u7fd2\u6163\u3068\u767d\u9aea\u306e\u65e9\u671f\u51fa\u73fe\u3068\u306e\u95a2\u9023\u304c\u5831\u544a\u3055\u308c\u3066\u304a\u308a\u3001\u7981\u7159\u306f\u767d\u9aea\u306e\u9032\u884c\u3092\u9045\u3089\u305b\u308b\u3046\u3048\u3067\u3082\u6709\u52b9\u306a\u53d6\u308a\u7d44\u307f\u3068\u3044\u3048\u307e\u3059\u3002<\/p>\n\n\n\n<h2 class=\"wp-block-heading\">\u6804\u990a\u4e0d\u8db3\u304c\u767d\u9aea\u3092\u52a0\u901f\u3055\u305b\u308b\uff5c\u9244\u30fb\u4e9c\u925b\u30fb\u30d3\u30bf\u30df\u30f3B12\u306f\u8db3\u308a\u3066\u3044\u307e\u3059\u304b<\/h2>\n\n\n\n<p class=\"wp-block-paragraph\">\u30e1\u30e9\u30cb\u30f3\u8272\u7d20\u3092\u6b63\u5e38\u306b\u5408\u6210\u3059\u308b\u305f\u3081\u306b\u306f\u3001\u7279\u5b9a\u306e\u30d3\u30bf\u30df\u30f3\u3084\u30df\u30cd\u30e9\u30eb\u304c\u6b20\u304b\u305b\u307e\u305b\u3093\u300240\u4ee3\u5973\u6027\u306f\u30c0\u30a4\u30a8\u30c3\u30c8\u3084\u504f\u98df\u3001\u6708\u7d4c\u306b\u3088\u308b\u9244\u640d\u5931\u306a\u3069\u304b\u3089\u6804\u990a\u4e0d\u8db3\u306b\u9665\u308a\u3084\u3059\u304f\u3001\u305d\u308c\u304c\u767d\u9aea\u306e\u9032\u884c\u3092\u65e9\u3081\u308b\u8981\u56e0\u3068\u306a\u3063\u3066\u3044\u307e\u3059\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u30d3\u30bf\u30df\u30f3B12\u4e0d\u8db3\u306f\u767d\u9aea\u306e\u5f15\u304d\u91d1\u306b\u306a\u308b<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\u30d3\u30bf\u30df\u30f3B12\u306fDNA\u5408\u6210\u3084\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u306e\u5897\u6b96\u306b\u95a2\u308f\u308b\u6804\u990a\u7d20\u3067\u3059\u3002\u3053\u306e\u30d3\u30bf\u30df\u30f3\u304c\u4e0d\u8db3\u3059\u308b\u3068\u3001\u6bdb\u5305\u306e\u6210\u9577\u671f\u304c\u4e0d\u5b89\u5b9a\u306b\u306a\u308a\u3001\u8272\u7d20\u7523\u751f\u304c\u4f4e\u4e0b\u3057\u3084\u3059\u304f\u306a\u308a\u307e\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u7814\u7a76\u3067\u306f\u3001\u767d\u9aea\u304c\u65e9\u671f\u306b\u51fa\u73fe\u3057\u305f\u60a3\u8005\u7fa4\u306b\u304a\u3044\u3066\u8840\u6e05\u30d3\u30bf\u30df\u30f3B12\u5024\u304c\u6709\u610f\u306b\u4f4e\u304b\u3063\u305f\u3053\u3068\u304c\u5831\u544a\u3055\u308c\u3066\u3044\u307e\u3059\u3002\u30d3\u30bf\u30df\u30f3B12\u306f\u8089\u985e\u30fb\u9b5a\u4ecb\u985e\u30fb\u4e73\u88fd\u54c1\u306b\u591a\u304f\u542b\u307e\u308c\u308b\u305f\u3081\u3001\u83dc\u98df\u4e2d\u5fc3\u306e\u98df\u751f\u6d3b\u3092\u9001\u3063\u3066\u3044\u308b\u65b9\u306f\u7279\u306b\u6ce8\u610f\u304c\u5fc5\u8981\u3067\u3057\u3087\u3046\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u9244\u3068\u9285\u306f\u30e1\u30e9\u30cb\u30f3\u751f\u6210\u306b\u6b20\u304b\u305b\u306a\u3044\u30df\u30cd\u30e9\u30eb<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\u9244\u306f\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u5185\u3067\u30e1\u30e9\u30cb\u30f3\u5408\u6210\u306b\u95a2\u308f\u308b\u9175\u7d20\u3092\u6d3b\u6027\u5316\u3059\u308b\u5f79\u5272\u3092\u62c5\u3044\u3001\u9285\u306f\u30c1\u30ed\u30b7\u30ca\u30fc\u30bc\u306e\u88dc\u56e0\u5b50\u3068\u3057\u3066\u50cd\u304d\u307e\u3059\u3002\u9285\u304c\u4e0d\u8db3\u3059\u308b\u3068\u30c1\u30ed\u30b7\u30ca\u30fc\u30bc\u306e\u6d3b\u6027\u304c\u4f4e\u4e0b\u3057\u3001\u30e6\u30fc\u30e1\u30e9\u30cb\u30f3\u3084\u30d5\u30a7\u30aa\u30e1\u30e9\u30cb\u30f3\u306e\u7523\u751f\u91cf\u304c\u843d\u3061\u308b\u3053\u3068\u304c\u308f\u304b\u3063\u3066\u3044\u307e\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u30a4\u30e9\u30f3\u306e\u7814\u7a76\u3067\u306f\u3001\u767d\u9aea\u306e\u3042\u308b\u60a3\u8005\u7fa4\u3067\u8840\u6e05\u9285\u6fc3\u5ea6\u304c\u5bfe\u7167\u7fa4\u3068\u6bd4\u3079\u3066\u6709\u610f\u306b\u4f4e\u3044\u3053\u3068\u304c\u793a\u3055\u308c\u307e\u3057\u305f\u3002\u666e\u6bb5\u306e\u98df\u4e8b\u304b\u3089\u8c9d\u985e\u3084\u30ca\u30c3\u30c4\u3001\u5927\u8c46\u88fd\u54c1\u3092\u610f\u8b58\u3057\u3066\u6442\u308b\u5de5\u592b\u304c\u3001\u9285\u306e\u88dc\u7d66\u306b\u52b9\u679c\u7684\u3067\u3059\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u30d3\u30bf\u30df\u30f3D\u3068\u30ab\u30eb\u30b7\u30a6\u30e0\u306e\u4e0d\u8db3\u3082\u898b\u843d\u3068\u305b\u306a\u3044<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\u30d3\u30bf\u30df\u30f3D\u306f\u30ab\u30eb\u30b7\u30a6\u30e0\u4ee3\u8b1d\u306b\u95a2\u4e0e\u3059\u308b\u3060\u3051\u3067\u306a\u304f\u3001\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u306e\u5206\u5316\u3084\u751f\u5b58\u306b\u3082\u5f71\u97ff\u3092\u4e0e\u3048\u308b\u3068\u3055\u308c\u3066\u3044\u307e\u3059\u3002\u65e5\u7167\u6642\u9593\u306e\u5c11\u306a\u3044\u5730\u57df\u306b\u4f4f\u3080\u65b9\u3084\u30aa\u30d5\u30a3\u30b9\u30ef\u30fc\u30af\u4e2d\u5fc3\u306e\u751f\u6d3b\u3092\u9001\u308b\u65b9\u306f\u3001\u30d3\u30bf\u30df\u30f3D\u304c\u4e0d\u8db3\u3057\u304c\u3061\u3067\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u5b66\u7ae5\u3092\u5bfe\u8c61\u3068\u3057\u305f\u7814\u7a76\u3067\u306f\u3042\u308a\u307e\u3059\u304c\u3001\u767d\u9aea\u306e\u3042\u308b\u88ab\u9a13\u8005\u7fa4\u3067\u30d3\u30bf\u30df\u30f3D\u6b20\u4e4f\u306e\u5272\u5408\u304c\u6709\u610f\u306b\u9ad8\u304b\u3063\u305f\u3068\u3044\u3046\u5831\u544a\u304c\u3042\u308a\u307e\u3059\u300240\u4ee3\u5973\u6027\u3082\u5ba4\u5185\u751f\u6d3b\u304c\u591a\u3044\u5834\u5408\u306f\u3001\u610f\u8b58\u7684\u306b\u65e5\u5149\u6d74\u3084\u30d3\u30bf\u30df\u30f3D\u542b\u6709\u98df\u54c1\u3092\u53d6\u308a\u5165\u308c\u308b\u3053\u3068\u304c\u5927\u5207\u3067\u3059\u3002<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>\u30d3\u30bf\u30df\u30f3B12\u3092\u542b\u3080\u98df\u54c1\uff1a\u30ec\u30d0\u30fc\u3001\u3042\u3055\u308a\u3001\u3057\u3058\u307f\u3001\u3055\u3093\u307e\u3001\u725b\u4e73<\/li>\n\n\n\n<li>\u9285\u3092\u542b\u3080\u98df\u54c1\uff1a\u7261\u8823\u3001\u30ab\u30b7\u30e5\u30fc\u30ca\u30c3\u30c4\u3001\u5927\u8c46\u3001\u30b3\u30b3\u30a2<\/li>\n\n\n\n<li>\u9244\u3092\u542b\u3080\u98df\u54c1\uff1a\u307b\u3046\u308c\u3093\u8349\u3001\u3072\u3058\u304d\u3001\u8d64\u8eab\u8089\u3001\u5c0f\u677e\u83dc<\/li>\n\n\n\n<li>\u30d3\u30bf\u30df\u30f3D\u3092\u542b\u3080\u98df\u54c1\uff1a\u9bad\u3001\u3057\u3089\u3059\u3001\u304d\u304f\u3089\u3052\u3001\u5375\u9ec4<\/li>\n<\/ul>\n\n\n\n<h2 class=\"wp-block-heading\">\u5fc3\u7406\u7684\u30b9\u30c8\u30ec\u30b9\u304c\u767d\u9aea\u3092\u5897\u3084\u3059\u306e\u306f\u79d1\u5b66\u7684\u306b\u8a3c\u660e\u3055\u308c\u3066\u3044\u308b<\/h2>\n\n\n\n<p class=\"wp-block-paragraph\">\u300c\u30b9\u30c8\u30ec\u30b9\u3067\u767d\u9aea\u304c\u5897\u3048\u305f\u300d\u3068\u3044\u3046\u8a71\u306f\u3088\u304f\u8033\u306b\u3057\u307e\u3059\u304c\u3001\u8fd1\u5e74\u306e\u7814\u7a76\u3067\u305d\u306e\u4ed5\u7d44\u307f\u304c\u79d1\u5b66\u7684\u306b\u88cf\u4ed8\u3051\u3089\u308c\u307e\u3057\u305f\u3002\u30b9\u30c8\u30ec\u30b9\u306f\u4ea4\u611f\u795e\u7d4c\u3092\u4ecb\u3057\u3066\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u5e79\u7d30\u80de\u3092\u67af\u6e07\u3055\u305b\u3001\u4e0d\u53ef\u9006\u7684\u306a\u767d\u9aea\u5316\u3092\u5f15\u304d\u8d77\u3053\u3059\u3053\u3068\u304c\u308f\u304b\u3063\u3066\u3044\u307e\u3059\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u4ea4\u611f\u795e\u7d4c\u306e\u904e\u5270\u306a\u6d3b\u6027\u5316\u304c\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u5e79\u7d30\u80de\u3092\u67af\u6e07\u3055\u305b\u308b<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">2020\u5e74\u306bNature\u8a8c\u3067\u767a\u8868\u3055\u308c\u305f\u30cf\u30fc\u30d0\u30fc\u30c9\u5927\u5b66\u306e\u7814\u7a76\u306f\u3001\u30b9\u30c8\u30ec\u30b9\u306b\u3088\u308b\u767d\u9aea\u5316\u306e\u4ed5\u7d44\u307f\u3092\u660e\u78ba\u306b\u793a\u3057\u307e\u3057\u305f\u3002\u6025\u6027\u30b9\u30c8\u30ec\u30b9\u3092\u53d7\u3051\u308b\u3068\u4ea4\u611f\u795e\u7d4c\u304c\u6d3b\u6027\u5316\u3057\u3001\u795e\u7d4c\u4f1d\u9054\u7269\u8cea\u30ce\u30eb\u30a2\u30c9\u30ec\u30ca\u30ea\u30f3\u304c\u5927\u91cf\u306b\u653e\u51fa\u3055\u308c\u307e\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u30ce\u30eb\u30a2\u30c9\u30ec\u30ca\u30ea\u30f3\u3092\u6d74\u3073\u305f\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u5e79\u7d30\u80de\u306f\u4f11\u7720\u72b6\u614b\u304b\u3089\u4e00\u6589\u306b\u76ee\u899a\u3081\u3001\u6025\u6fc0\u306b\u5897\u6b96\u30fb\u5206\u5316\u3057\u3066\u3057\u307e\u3044\u307e\u3059\u3002\u672c\u6765\u3086\u3063\u304f\u308a\u3068\u8272\u7d20\u3092\u4f9b\u7d66\u3059\u308b\u306f\u305a\u306e\u5e79\u7d30\u80de\u304c\u4e00\u5ea6\u306b\u4f7f\u3044\u679c\u305f\u3055\u308c\u308b\u3053\u3068\u3067\u3001\u6bdb\u5305\u306f\u8272\u7d20\u3092\u4f5c\u308b\u80fd\u529b\u3092\u6c38\u4e45\u306b\u5931\u3046\u306e\u3067\u3059\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u30b9\u30c8\u30ec\u30b9\u306b\u3088\u308b\u767d\u9aea\u306f\u6761\u4ef6\u6b21\u7b2c\u3067\u56de\u5fa9\u3067\u304d\u308b<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\u4e00\u65b9\u3067\u30012021\u5e74\u306beLife\u8a8c\u3067\u5831\u544a\u3055\u308c\u305f\u7814\u7a76\u3067\u306f\u3001\u30b9\u30c8\u30ec\u30b9\u304c\u89e3\u6d88\u3055\u308c\u308b\u3053\u3068\u3067\u4e00\u90e8\u306e\u767d\u9aea\u304c\u81ea\u7136\u306b\u518d\u8272\u7d20\u5316\uff08\u3082\u3068\u306e\u8272\u306b\u623b\u308b\uff09\u3059\u308b\u30b1\u30fc\u30b9\u304c\u78ba\u8a8d\u3055\u308c\u307e\u3057\u305f\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u88ab\u9a13\u8005\u306e\u30b9\u30c8\u30ec\u30b9\u65e5\u8a18\u3068\u6bdb\u9aea\u306e\u8272\u7d20\u30d1\u30bf\u30fc\u30f3\u3092\u7167\u5408\u3057\u305f\u7d50\u679c\u3001\u30b9\u30c8\u30ec\u30b9\u306e\u8efd\u6e1b\u3068\u767d\u9aea\u306e\u56de\u5fa9\u304c\u6642\u9593\u7684\u306b\u4e00\u81f4\u3057\u3066\u3044\u305f\u306e\u3067\u3059\u3002<\/p>\n\n\n\n<h4 class=\"wp-block-heading\">\u30b9\u30c8\u30ec\u30b9\u3068\u767d\u9aea\u306e\u95a2\u4fc2<\/h4>\n\n\n\n<figure class=\"wp-block-table\"><table class=\"has-fixed-layout\"><thead><tr><th>\u72b6\u614b<\/th><th>\u30e1\u30e9\u30ce\u30b5\u30a4\u30c8\u5e79\u7d30\u80de\u3078\u306e\u5f71\u97ff<\/th><th>\u767d\u9aea\u306e\u56de\u5fa9\u53ef\u80fd\u6027<\/th><\/tr><\/thead><tbody><tr><td>\u6025\u6027\u30b9\u30c8\u30ec\u30b9<\/td><td>\u5e79\u7d30\u80de\u306e\u6025\u901f\u306a\u67af\u6e07<\/td><td>\u6761\u4ef6\u306b\u3088\u308a\u4e00\u90e8\u56de\u5fa9<\/td><\/tr><tr><td>\u6162\u6027\u30b9\u30c8\u30ec\u30b9<\/td><td>\u5e79\u7d30\u80de\u306e\u6301\u7d9a\u7684\u306a\u6d88\u8017<\/td><td>\u56de\u5fa9\u306f\u56f0\u96e3\u306a\u5834\u5408\u304c\u591a\u3044<\/td><\/tr><tr><td>\u30b9\u30c8\u30ec\u30b9\u89e3\u6d88\u5f8c<\/td><td>\u6b8b\u5b58\u5e79\u7d30\u80de\u304c\u518d\u6d3b\u6027\u5316<\/td><td>\u5e79\u7d30\u80de\u304c\u6b8b\u3063\u3066\u3044\u308c\u3070\u53ef\u80fd<\/td><\/tr><\/tbody><\/table><\/figure>\n\n\n\n<h3 class=\"wp-block-heading\">40\u4ee3\u5973\u6027\u306b\u30b9\u30c8\u30ec\u30b9\u6027\u767d\u9aea\u304c\u591a\u3044\u80cc\u666f<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">40\u4ee3\u306f\u4ed5\u4e8b\u3067\u306e\u8cac\u4efb\u304c\u5897\u3059\u6642\u671f\u3067\u3042\u308b\u3068\u3068\u3082\u306b\u3001\u5b50\u80b2\u3066\u3084\u89aa\u306e\u4ecb\u8b77\u304c\u91cd\u306a\u308a\u3084\u3059\u3044\u5e74\u4ee3\u3067\u3082\u3042\u308a\u307e\u3059\u3002\u30db\u30eb\u30e2\u30f3\u30d0\u30e9\u30f3\u30b9\u306e\u4e71\u308c\u3082\u52a0\u308f\u308b\u305f\u3081\u3001\u5fc3\u8eab\u306e\u30b9\u30c8\u30ec\u30b9\u304c\u8907\u5408\u7684\u306b\u6bdb\u6839\u3078\u5f71\u97ff\u3092\u4e0e\u3048\u3084\u3059\u3044\u306e\u3067\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u767d\u9aea\u3092\u6c17\u306b\u3059\u308b\u3053\u3068\u81ea\u4f53\u304c\u307e\u305f\u65b0\u305f\u306a\u30b9\u30c8\u30ec\u30b9\u306b\u306a\u308b\u3068\u3044\u3046\u60aa\u5faa\u74b0\u306b\u9665\u308b\u30b1\u30fc\u30b9\u3082\u3042\u308b\u304b\u3082\u3057\u308c\u307e\u305b\u3093\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u5b8c\u74a7\u3092\u76ee\u6307\u3059\u306e\u3067\u306f\u306a\u304f\u3001\u3067\u304d\u308b\u7bc4\u56f2\u3067\u30b9\u30c8\u30ec\u30b9\u3092\u6e1b\u3089\u3059\u5de5\u592b\u3092\u53d6\u308a\u5165\u308c\u308b\u3053\u3068\u304c\u3001\u767d\u9aea\u306e\u9032\u884c\u3092\u7de9\u3084\u304b\u306b\u3059\u308b\u7b2c\u4e00\u6b69\u306b\u306a\u308a\u307e\u3059\u3002<\/p>\n\n\n\n<h2 class=\"wp-block-heading\">\u3053\u308c\u4ee5\u4e0a\u767d\u9aea\u3092\u5897\u3084\u3055\u306a\u3044\u305f\u3081\u306b\u4eca\u65e5\u304b\u3089\u5909\u3048\u305f\u3044\u751f\u6d3b\u7fd2\u6163<\/h2>\n\n\n\n<p 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class=\"wp-block-heading\">\u982d\u76ae\u30de\u30c3\u30b5\u30fc\u30b8\u3067\u8840\u6d41\u3092\u4fc3\u9032\u3059\u308b<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\u982d\u76ae\u306e\u8840\u884c\u304c\u60aa\u304f\u306a\u308b\u3068\u3001\u6bdb\u6839\u306b\u5c4a\u304f\u9178\u7d20\u3084\u6804\u990a\u7d20\u304c\u4e0d\u8db3\u3057\u304c\u3061\u3067\u3059\u3002\u5165\u6d74\u6642\u3084\u30b7\u30e3\u30f3\u30d7\u30fc\u6642\u306b\u6307\u306e\u8179\u3067\u982d\u76ae\u3092\u3084\u3055\u3057\u304f\u63c9\u307f\u307b\u3050\u3059\u30de\u30c3\u30b5\u30fc\u30b8\u3092\u7fd2\u6163\u306b\u3059\u308b\u3068\u3001\u8840\u6d41\u6539\u5584\u304c\u671f\u5f85\u3067\u304d\u307e\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u5f37\u304f\u3053\u3059\u308a\u3059\u304e\u308b\u3068\u982d\u76ae\u3092\u50b7\u3081\u3066\u3057\u307e\u3046\u305f\u3081\u3001\u6c17\u6301\u3061\u3088\u3044\u3068\u611f\u3058\u308b\u7a0b\u5ea6\u306e\u529b\u52a0\u6e1b\u304c\u9069\u5207\u3067\u3059\u30021\u56de3\u301c5\u5206\u307b\u3069\u306e\u77ed\u3044\u6642\u9593\u3067\u3082\u3001\u6bce\u65e5\u7d9a\u3051\u308b\u3068\u982d\u76ae\u74b0\u5883\u306b\u5909\u5316\u304c\u51fa\u3084\u3059\u304f\u306a\u308a\u307e\u3059\u3002<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>\u7981\u7159\u306f\u9178\u5316\u30b9\u30c8\u30ec\u30b9\u306e\u4f4e\u6e1b\u306b\u76f4\u7d50\u3059\u308b<\/li>\n\n\n\n<li>\u904e\u5ea6\u306a\u30c0\u30a4\u30a8\u30c3\u30c8\u306f\u6804\u990a\u4e0d\u8db3\u3092\u62db\u304d\u767d\u9aea\u3092\u4fc3\u9032\u3059\u308b<\/li>\n\n\n\n<li>\u30a6\u30a9\u30fc\u30ad\u30f3\u30b0\u3084\u30e8\u30ac\u306a\u3069\u9069\u5ea6\u306a\u6709\u9178\u7d20\u904b\u52d5\u306f\u8840\u884c\u3092\u6539\u5584\u3059\u308b<\/li>\n\n\n\n<li>\u98f2\u9152\u91cf\u3092\u898b\u76f4\u3059\u3053\u3068\u3082\u6bdb\u6839\u306e\u5065\u5eb7\u7dad\u6301\u306b\u5bc4\u4e0e\u3059\u308b<\/li>\n<\/ul>\n\n\n\n<h2 class=\"wp-block-heading\">\u767d\u9aea\u67d3\u3081\u3060\u3051\u306b\u983c\u3089\u306a\u3044\uff5c\u982d\u76ae\u30b1\u30a2\u3068\u533b\u7642\u6a5f\u95a2\u3067\u306e\u76f8\u8ac7<\/h2>\n\n\n\n<p class=\"wp-block-paragraph\">\u767d\u9aea\u304c\u76ee\u7acb\u3061\u59cb\u3081\u308b\u3068\u767d\u9aea\u67d3\u3081\u306b\u983c\u308a\u304c\u3061\u3067\u3059\u304c\u3001\u982d\u76ae\u3078\u306e\u30c0\u30e1\u30fc\u30b8\u3084\u67d3\u3081\u76f4\u3057\u306e\u624b\u9593\u3092\u8003\u3048\u308b\u3068\u3001\u305d\u308c\u3060\u3051\u3067\u306f\u6839\u672c\u7684\u306a\u89e3\u6c7a\u306b\u306f\u306a\u308a\u307e\u305b\u3093\u3002\u982d\u76ae\u30b1\u30a2\u306e\u898b\u76f4\u3057\u3084\u3001\u5fc5\u8981\u306b\u5fdc\u3058\u3066\u533b\u7642\u6a5f\u95a2\u3078\u76f8\u8ac7\u3059\u308b\u3053\u3068\u3082\u9078\u629e\u80a2\u306b\u5165\u308c\u3066\u307f\u3066\u304f\u3060\u3055\u3044\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u767d\u9aea\u67d3\u3081\u3092\u7d9a\u3051\u308b\u3068\u304d\u982d\u76ae\u30c0\u30e1\u30fc\u30b8\u3092\u6700\u5c0f\u9650\u306b\u6291\u3048\u308b\u65b9\u6cd5<\/h3>\n\n\n\n<p 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class=\"has-fixed-layout\"><thead><tr><th>\u7a2e\u985e<\/th><th>\u6301\u7d9a\u671f\u9593<\/th><th>\u982d\u76ae\u3078\u306e\u8ca0\u62c5<\/th><\/tr><\/thead><tbody><tr><td>\u6c38\u4e45\u67d3\u6bdb\u5264<\/td><td>1\u301c2\u304b\u6708\u7a0b\u5ea6<\/td><td>\u6bd4\u8f03\u7684\u9ad8\u3044<\/td><\/tr><tr><td>\u534a\u6c38\u4e45\u67d3\u6bdb\u6599<\/td><td>2\u301c4\u9031\u9593\u7a0b\u5ea6<\/td><td>\u4e2d\u7a0b\u5ea6<\/td><\/tr><tr><td>\u4e00\u6642\u67d3\u6bdb\u6599<\/td><td>\u30b7\u30e3\u30f3\u30d7\u30fc\u3067\u843d\u3061\u308b<\/td><td>\u4f4e\u3044<\/td><\/tr><tr><td>\u30d8\u30ca\uff08\u5929\u7136\u67d3\u6599\uff09<\/td><td>3\u301c4\u9031\u9593\u7a0b\u5ea6<\/td><td>\u6bd4\u8f03\u7684\u4f4e\u3044<\/td><\/tr><\/tbody><\/table><\/figure>\n\n\n\n<h3 class=\"wp-block-heading\">\u5e02\u8ca9\u306e\u767d\u9aea\u5bfe\u7b56\u30b5\u30d7\u30ea\u30e1\u30f3\u30c8\u306f\u52b9\u304f\u306e\u304b<\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\u30d3\u30bf\u30df\u30f3B12\u3001\u30d3\u30aa\u30c1\u30f3\u3001\u9285\u3001\u4e9c\u925b\u306a\u3069\u3092\u914d\u5408\u3057\u305f\u30b5\u30d7\u30ea\u30e1\u30f3\u30c8\u304c\u767d\u9aea\u5bfe\u7b56\u3068\u3057\u3066\u8ca9\u58f2\u3055\u308c\u3066\u3044\u307e\u3059\u304c\u3001\u73fe\u6642\u70b9\u3067\u306f\u30b5\u30d7\u30ea\u30e1\u30f3\u30c8\u3060\u3051\u3067\u767d\u9aea\u3092\u4e88\u9632\u30fb\u6539\u5584\u3067\u304d\u308b\u3068\u3044\u3046\u5341\u5206\u306a\u79d1\u5b66\u7684\u6839\u62e0\u306f\u3042\u308a\u307e\u305b\u3093\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u305f\u3060\u3057\u3001\u8840\u6db2\u691c\u67fb\u3067\u7279\u5b9a\u306e\u6804\u990a\u7d20\u306e\u6b20\u4e4f\u304c\u78ba\u8a8d\u3055\u308c\u305f\u5834\u5408\u306b\u306f\u3001\u4e0d\u8db3\u5206\u3092\u88dc\u3046\u3053\u3068\u3067\u767d\u9aea\u306e\u9032\u884c\u304c\u7de9\u3084\u304b\u306b\u306a\u308b\u53ef\u80fd\u6027\u306f\u3042\u308a\u307e\u3059\u3002<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\u81ea\u5df1\u5224\u65ad\u3067\u5927\u91cf\u306b\u6442\u53d6\u3059\u308b\u306e\u3067\u306f\u306a\u304f\u3001\u533b\u5e2b\u3084\u6804\u990a\u58eb\u306b\u76f8\u8ac7\u3057\u305f\u3046\u3048\u3067\u9069\u5207\u306a\u91cf\u3092\u88dc\u7d66\u3059\u308b\u3053\u3068\u3092\u304a\u3059\u3059\u3081\u3057\u307e\u3059\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\">\u533b\u7642\u6a5f\u95a2\u3067\u53d7\u3051\u3089\u308c\u308b\u767d\u9aea\u95a2\u9023\u306e\u8a3a\u5bdf\u5185\u5bb9<\/h3>\n\n\n\n<p 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class=\"swell-block-accordion\">\n<details class=\"swell-block-accordion__item\" data-swl-acc=\"wrapper\"><summary class=\"swell-block-accordion__title\" data-swl-acc=\"header\"><span class=\"swell-block-accordion__label\">\u53c2\u8003\u6587\u732e<\/span><span class=\"swell-block-accordion__icon c-switchIconBtn\" data-swl-acc=\"icon\" aria-hidden=\"true\" data-opened=\"false\"><i class=\"__icon--closed icon-caret-down\"><\/i><i class=\"__icon--opened icon-caret-up\"><\/i><\/span><\/summary><div class=\"swell-block-accordion__body\" data-swl-acc=\"body\">\n<p class=\"wp-block-paragraph\">Wood, J. M., Decker, H., Hartmann, H., Chavan, B., Rokos, H., Spencer, J. D., Hasse, S., Thornton, M. J., Shalbaf, M., Paus, R., &amp; Schallreuter, K. U. (2009). Senile hair graying: H2O2-mediated oxidative stress affects human hair color by blunting methionine sulfoxide repair. FASEB Journal, 23(7), 2065\u20132075. https:\/\/doi.org\/10.1096\/fj.08-125435<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Desai, D. D., Karim, M., Nohria, A., Needle, C., Brinks, A., Kearney, C. A., Ridge, A., Mesinkovska, N., Shapiro, J., &amp; Lo Sicco, K. I. (2025). Premature hair graying: A multifaceted phenomenon. International Journal of Dermatology, 64(5), 819\u2013829. https:\/\/doi.org\/10.1111\/ijd.17580<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Rosenberg, A. M., Rausser, S., Ren, J., Mosharov, E. V., Sturm, G., Ogden, R. T., Patel, P., Soni, R. K., Lacefield, C., Tobin, D. J., Paus, R., &amp; Picard, M. (2021). Quantitative mapping of human hair greying and reversal in relation to life stress. eLife, 10, e67437. https:\/\/doi.org\/10.7554\/eLife.67437<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Sun, Q., Lee, W., Hu, H., Ogawa, T., De Leon, S., Katehis, I., Lim, C. H., Takeo, M., Cammer, M., Taketo, M. M., Gay, D. L., Millar, S. E., &amp; Ito, M. (2023). Dedifferentiation maintains melanocyte stem cells in a dynamic niche. Nature, 616(7958), 774\u2013782. https:\/\/doi.org\/10.1038\/s41586-023-05960-6<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">O&#8217;Sullivan, J. D. B., Nicu, C., Picard, M., Ch\u00e9ret, J., Bedogni, B., Tobin, D. J., &amp; Paus, R. (2021). The biology of human hair greying. Biological Reviews, 96(1), 107\u2013128. https:\/\/doi.org\/10.1111\/brv.12648<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Nishimura, E. K., Granter, S. R., &amp; Fisher, D. E. (2005). Mechanisms of hair graying: Incomplete melanocyte stem cell maintenance in the niche. Science, 307(5710), 720\u2013724. https:\/\/doi.org\/10.1126\/science.1099593<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Zhang, B., Ma, S., Rachmin, I., He, M., Baral, P., Choi, S., Gon\u00e7alves, W. A., Shwartz, Y., Fast, E. M., Su, Y., Zon, L. I., Regev, A., Buenrostro, J. D., Cunha, T. M., Chiu, I. M., Fisher, D. E., &amp; Hsu, Y.-C. (2020). Hyperactivation of sympathetic nerves drives depletion of melanocyte stem cells. Nature, 577(7792), 676\u2013681. https:\/\/doi.org\/10.1038\/s41586-020-1935-3<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Kumar, A. B., Shamim, H., &amp; Nagaraju, U. (2018). Premature graying of hair: Review with updates. 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